| Category Name |
Traits Name |
Traits Description |
| Life history |
Adult |
Identifies if traits were compiled for aquatic adults, otherwise entries pertain to immature life stage (entries: ADULT or blank) |
| Life history |
No. of Aquatic stages |
Number of aquatic life stages (4 potential entries) |
| Life history - development |
Ability to survive abbreviated |
Abbreviated ability to survive desiccation entries: present = able to survive desiccation; absent = not able to survive desiccation |
| Life history - development |
Ability to survive comments |
Description of abbreviated adult ability to survive desiccation entries: present = able to survive desiccation; absent = not able to survive desiccation |
| Life history - development |
Adult ability to temporarily exit water |
Indicates ability to temporarily exit water. Entries = no, yes, unknown, blank |
| Life history - development |
Adult ability to temporarily exit water abbreviated |
Abbreviated adult ability to exit entries: present = has ability to exit; absent = does NOT have ability to exit |
| Life history - development |
Adult ability to temporarily exit water comments |
Description of abbreviated adult ability to exit entries: present = has ability to exit; absent = does NOT have ability to exit. This does NOT include emergence. |
| Life history - development |
Development pattern comments |
Development pattern text notes |
| Life history - development |
Development speed |
Development speed. Entries: fast seasonal, slow seasonal, Non-seasonal |
| Life history - development |
Development speed abbreviated |
Abbreviated development entries: fast, slow, non |
| Life history - development |
Diapause |
Indicates whether diapause occurs. Entries: no, yes, unknown, blank |
| Life history - development |
Hatch time |
Time required for eggs to hatch. Entries: hours, minutes, days, months, weeks |
| Life history - development |
Hatch time comments |
Time required for eggs to hatch text notes |
| Life history - development |
Life span |
Adult lifespan. Entries: days, hours, weeks, months |
| Life history - development |
Life span abbreviated |
Abbreviated adult life span: very_short, short, long |
| Life history - development |
Life span comments |
Description of abbreviated adult life span entries: very short = less than 1 week; short = less than 1 month; long = greater than 1 month |
| Life history - emergence |
Duration of oviposition period |
Duration of oviposition period. Entries = days, months, weeks |
| Life history - emergence |
Egg - mass |
Egg type - one mass |
| Life history - emergence |
Egg - mutiple batches |
Egg type - multiple batches |
| Life history - emergence |
Egg - single |
Egg type - Single |
| Life history - emergence |
Egg cemented |
Indicates whether eggs are cemented. Entries = no, yes |
| Life history - emergence |
Emergence begins |
Season that emergence begins |
| Life history - emergence |
Emergence behavior - drifting |
emergence behavior - drifting |
| Life history - emergence |
Emergence behavior comments |
Emergence behavior text notes |
| Life history - emergence |
Emergence comments |
Seasons during which sexually mature forms have been reported. Entries = winter, spring, summer, fall |
| Life history - emergence |
Emergence ends |
Season that emergence ends |
| Life history - emergence |
Emergence occur all year |
Indicates whether emergence can occur all year. Entries = no, yes, unknown, blank |
| Life history - emergence |
Emergence synchronization |
Indicates whether emergence is synchronous. Entries = no, yes, unknown, blank |
| Life history - emergence |
Emergence synchronization Abbreviated |
Abbreviated synchronization of emergence entries = poorly, well |
| Life history - emergence |
Emergence synchronization comments |
Description of abbreviated synchronization of emergence entries: poorly = week; well = days |
| Life history - emergence |
Fecundity |
Fecundity. Entries: < 100 eggs, > 10,000 eggs, 100 to 1000 eggs, 1000 to 10,000 eggs |
| Life history - emergence |
Oviposition behavior comments |
Oviposition behavior text notes |
| Life history - emergence |
Primary oviposition behavior |
Primary oviposition behavior (10-15 potential entries) |
| Life history - emergence |
Secondary oviposition behavior |
Secondary oviposition behavior (10-15 potential entries) |
| Life history - emergence |
Voltinism |
Voltinism. Entries: > 1 Generation per year, 1 Generation per year, < 1 Generation per year |
| Life history - emergence |
Voltinism Comments |
Voltinism text comments (i.e., overwintering of eggs or immatures) |
| Life history - emergence |
Voltinism abbreviated |
Abbreviated voltinism entries: semivoltine, univoltine, bi_multivoltine |
| Life history - emergence |
emergence behavior - climbing |
emergence behavior - climbing |
| Life history - emergence |
emergence behavior - crawling |
emergence behavior - crawling |
| Mobility |
Adult dispersal distance |
Adult dispersal distance. Entries = 1 km or less, 10 km or less, 10 m or less, 100 km or less |
| Mobility |
Drift abbreviated |
Abbreviated occurrence in drift entries: rare, common, abundant |
| Mobility |
Drift propensity comments |
Description of abbreviated occurrence in drift entries: rare = catastrophic only; common = typically observed; abundant = dominant in drift samples |
| Mobility |
Drift propensity of early instars |
Drift propensity of early instars. Entries = Strong (active / often), Medium (mostly passive / occasional), Weak (catastrophic only) |
| Mobility |
Drift propensity of late instars |
Drift propensity of late instars. Entries = Strong (active / often), Medium (mostly passive / occasional), Weak (catastrophic only) |
| Mobility |
Female dispersal |
Abbreviated female dispersal entries: low, high |
| Mobility |
Female dispersal comments |
Descripton of abbreviated female dispersal entries: low = less than 1 km flight before laying eggs; high = greater than 1 km flight before laying eggs |
| Mobility |
Flying strength |
Abbreviated flying strength entries: weak, strong |
| Mobility |
Flying strength comments |
Description of abbreviated flying strength entries: weak = e.g., cannot fly into light breeze |
| Mobility |
Larval dispersal distance |
Larval dispersal distance. Entries = < 1 m, 1-10 m, 11-100 m |
| Mobility |
Maximum crawling rate |
Abbreviated maximum crawling rate entries: very_low, low, high |
| Mobility |
Maximum crawling rate comments |
Description of abbreviated maximum crawling rate entries: very low = less than 10 cm per hour; low = less than 100 cm per hour; high = greater than 100 cm per hour |
| Mobility |
Swimming ability |
Abbreviated swimming ability entries: none, weak strong |
| Morphology - Adaptation |
Ballast |
taxon has ballast |
| Morphology - Adaptation |
Friction |
taxon has friction pads or other structures to reduce friction coefficient with surface |
| Morphology - Adaptation |
Hair |
taxon has hair |
| Morphology - Adaptation |
Hooks |
taxon has hooks |
| Morphology - Adaptation |
Other |
Text to further describe morphological adaptations |
| Morphology - Adaptation |
Silk |
taxon has silk |
| Morphology - Adaptation |
Suckers |
taxon has suckers |
| Morphology - Armor |
Armor |
Degree of body armoring. Entries = All sclerotized, Hard shelled, Partly sclerotized, Soft |
| Morphology - Armor |
Armor abbreviated |
Abbreviated armoring entries = none, poor, good |
| Morphology - Armor |
Armor comments |
Description of abbreviated armoring entries: none = soft-bodied forms; poor = heavily sclerotized; good = e.g., some cased caddisflies |
| Morphology - Attachment |
Attachment abbreviated |
Abbreviated attachment entries = none, some, both |
| Morphology - Attachment |
Attachment comments |
Abbreviated attachment entries: none = free-ranging; some = sessile, sedentary; both = free-ranging and sessile, sedentary. Other (non-abbreviated) entries include: normally free living and capable of locomotion; both sessile and free living and capable of locomotion; normally sessile. |
| Morphology - Body |
Body Shape abbreviated |
Abbreviated body shape entries = streamlined, not_streamlined |
| Morphology - Body |
Body height |
Measured body height of immatures (mm) |
| Morphology - Body |
Body length |
Measured body length of immatures (mm) |
| Morphology - Body |
Body shape |
Body shape. Entries = Bluff (blocky), Dorsoventrally flattened, Round (humped), Streamlined / fusiform, Tubular |
| Morphology - Body |
Body shape case |
Body shape with case/retreat. Entries = Bluff (blocky), Dorsoventrally flattened, Round (humped), Streamlined / fusiform, Tubular |
| Morphology - Body |
Body shape comments |
Description of abbreviated body shape entries: streamlined = flat, fusiform; not streamlined = cylindrical, round or bluff |
| Morphology - Body |
Body width |
Measured body width of immatures (mm) |
| Morphology - Body |
Maximal body size |
Maximal body size of immatures. Entries = Large (length > 16 mm), Medium (length 9-16 mm), Small (length < 9 mm) |
| Morphology - Body |
Mediate drag |
Indication of whether shape mediates drag. Entries = no, yes, unknown, blank |
| Morphology - Respiration |
Respiration abbreviated |
Abbreviated respiration entries = tegument, gills, plastron_spiracle |
| Morphology - Respiration |
Respiration comments |
Respiration text notes |
| Morphology - Respiration |
Respiration mode of aquatic adults |
Respiration mode of aquatic adults |
| Morphology - Respiration |
Respiration mode of early instars |
Respiration mode of early instars |
| Morphology - Respiration |
Respiration mode of late instars |
Respiration mode of late instars |
| Resource Acquisition Preference |
Current quiet |
current preference - quiet |
| Resource Acquisition Preference |
Current slow |
current preference - slow |
| Resource Acquisition Preference |
Feed mode comments |
Description of abbreviated primary functional feeding group entries: CF = collector-filterer; CG = collector-gatherer; HB = herbivore (scraper); SH = shredder; PR = predator (piercer, engulfer); PA = parasite. Other (non-abbreviated) entries include text notes on food material consumed. |
| Resource Acquisition Preference |
Habit abbreviated |
Abbreviated habit entries: BU, CB, CN, SK, SP, SW |
| Resource Acquisition Preference |
Habit comments |
Description of abbreviated primary habit entries: BU = burrower; CB = climber; CN = clinger; SK = skater; SP = sprawler; SW = swimmer. Other (non-abbreviated) entries include text notes on habit. |
| Resource Acquisition Preference |
Lateral habitat position comments |
Lateral habitat position in water column text notes |
| Resource Acquisition Preference |
Lateral habitat position hyporheic |
Lateral habitat position in water column - hyporheic |
| Resource Acquisition Preference |
Lateral habitat position margin |
Lateral habitat position in water column - margin |
| Resource Acquisition Preference |
Lateral habitat position pool |
Lateral habitat position in water column - pool |
| Resource Acquisition Preference |
Lateral habitat position riffle |
Lateral habitat position in water column - riffle |
| Resource Acquisition Preference |
Lateral habitat position shoreline |
Lateral habitat position in water column - shoreline |
| Resource Acquisition Preference |
Microhabitat algae |
Microhabitat substrate preference - algae |
| Resource Acquisition Preference |
Microhabitat boulder |
Microhabitat substrate preference - boulder |
| Resource Acquisition Preference |
Microhabitat comments |
Microhabitat substrate preference text notes |
| Resource Acquisition Preference |
Microhabitat detritus |
Microhabitat substrate preference - detritus |
| Resource Acquisition Preference |
Microhabitat gravel |
Microhabitat substrate preference - gravel |
| Resource Acquisition Preference |
Microhabitat large woody debris |
Microhabitat substrate preference - large woody debris (LWD) |
| Resource Acquisition Preference |
Microhabitat macrophytes |
Microhabitat substrate preference - macrophytes |
| Resource Acquisition Preference |
Microhabitat pelagic |
Microhabitat substrate preference - pelagic |
| Resource Acquisition Preference |
Microhabitat plants |
Microhabitat substrate preference - plants |
| Resource Acquisition Preference |
Microhabitat rocks |
Microhabitat substrate preference - rocks |
| Resource Acquisition Preference |
Microhabitat sand |
Microhabitat substrate preference - sand |
| Resource Acquisition Preference |
Microhabitat silt |
Microhabitat substrate preference - silt |
| Resource Acquisition Preference |
Primary feeding mode |
Primary feeding mode based on mouthpart morphology |
| Resource Acquisition Preference |
Primary functional feeding group abbreviated |
Abbreviated primary functional feeding group entries: CF, CG, HB, PA, PR, SH |
| Resource Acquisition Preference |
Primary habit |
Primary habit |
| Resource Acquisition Preference |
Rheophily |
Abbreviated rheophily entries: depo, depo_eros, eros |
| Resource Acquisition Preference |
Rheophily comments |
Description of abbreviated rheophily entries: depo = depositional only, depo_eros = depositional and erosional; eros: erosional only. Other (categorical) entries include: fast, moderate, moderate-fast, standing and flowing, standing-slight, slight. |
| Resource Acquisition Preference |
Secondary feeding mode |
Secondary feeding mode based on mouthpart morphology |
| Resource Acquisition Preference |
Secondary habit |
Secondary habit |
| Resource Acquisition Preference |
Vertical habitat position benthic |
Vertical habitat position in water column - benthic |
| Resource Acquisition Preference |
Vertical habitat position comments |
Vertical habitat position in water column text notes |
| Resource Acquisition Preference |
Vertical habitat position hyporheic |
Vertical habitat position in water column - hyporheic |
| Resource Acquisition Preference |
Vertical habitat position macrophytes |
Vertical habitat position in water column - macrophytes |
| Resource Acquisition Preference |
Vertical habitat position pelagic |
Vertical habitat position in water column - pelagic |
| Resource Acquisition Preference |
Vertical habitat position surface |
Vertical habitat position in water column - surface |
| Resource Acquisition Preference |
current fast laminar |
current preference - fast laminar currents |
| Resource Acquisition Preference |
current fast turbulent |
current preference - fast turbulent currents |
| Resource Acquisition Preference |
current moderate |
current preference - moderate |
| Tolerance |
Current optima rank |
Rank values were derived using an arbitrary 1-7 scoring scheme, based on percentiles within the data set. Low scores = preference for slower water and high scores = preference for faster water. For more information, see Data Source Summary document. |
| Tolerance |
Current optima value |
Optima values for current (velocity) data (typically derived from weighted average or generalized linear model inferences). For more information, see Data Source Summary document. |
| Tolerance |
Enrichment tolerance score |
Numerical tolerance score ranging from 0 (most intolerant) to 10 (most tolerant). Typically based on tolerances to organic enrichment. |
| Tolerance |
Enrichment tolerance score comments |
Description of enrichment tolerance scores and sources |
| Tolerance |
Eurythermal |
Taxon documented in eurythermal (< 15 C) temperature range |
| Tolerance |
Euthermal |
Taxon documented in euthermal (< 30 C) temperature range |
| Tolerance |
High DO levels |
Oxygen tolerance - high DO levels |
| Tolerance |
Low DO levels |
Oxygen tolerance - low DO levels |
| Tolerance |
Maximum temperature reported |
Maximum temperature reported |
| Tolerance |
Mesothermal |
Taxon documented in mesothermal (15-30 C) temperature range |
| Tolerance |
Metathermal |
Taxon documented in metathermal (5-15 C) temperature range |
| Tolerance |
Minimum temperature reported |
Minimum temperature reported |
| Tolerance |
Normal DO levels |
Oxygen tolerance - normal (intermediate) DO levels |
| Tolerance |
Observed lethal DO levels |
Observed lethal DO levels |
| Tolerance |
Observed maximum letal temperature |
Observed maximum letal temperature |
| Tolerance |
Oligothermal |
Taxon documented in oligothermal (< 15 C) temperature range |
| Tolerance |
Oxygen tolerance comments |
General oxygen tolerance categories: High, Moderate, Moderate-High, Low, Low-Moderate, Anaerobic, Low-Anaerobic, No strong preference |
| Tolerance |
Ph comments |
General pH tolerance categories: Acidic, Acid-Neutral, Alkaline, Alkaline-Neutral, Neutral, No strong preference |
| Tolerance |
Salinity tolerance - brakish |
Salinity tolerance - brakish |
| Tolerance |
Salinity tolerance - fresh |
Salinity tolerance - fresh |
| Tolerance |
Salinity tolerance - saline |
Salinity tolerance - saline |
| Tolerance |
Stenothermal |
Taxon documented in stenothermal (< 5 C) temperature range |
| Tolerance |
Thermal Indicator |
Cold and warm water preference taxa for particular states or regions. Entries = cold, warm, blank. NOTE: these lists are preliminary. For more information, see Data Source Summary document. |
| Tolerance |
Thermal comments |
Text notes pertaining to thermal entries. |
| Tolerance |
Thermal optima rank |
Rank values were derived using an arbitrary 1-7 scoring scheme, based on percentiles within the data set. Low scores = preference for colder water and high scores = preference for warmer water. For more information, see Data Source Summary document. |
| Tolerance |
Thermal optima value |
Optima values for temperature data (typically derived from weighted average or generalized linear model inferences). For more information, see Data Source Summary document. |
| Tolerance |
Thermal preference categories |
General thermal preference categories: Cold stenothermal (<5 C), Cold-cool eurythermal (0-15 C), Warm eurythermal (15-30 C), Hot euthermal (>30 C), No strong preference |
| Tolerance |
Thermal tolerance rank |
Rank values were derived using an arbitrary 1-7 scoring scheme, based on percentiles within the data set. Low scores = narrow thermal tolerance range and high scores = wide thermal tolerance range. For more information, see Data Source Summary document. |
| Tolerance |
Thermal tolerance value |
Tolerance values for temperature data (typically derived from weighted average or generalized linear model inferences). For more information, see Data Source Summary document. |
| Tolerance |
Turbidity |
General turbidity tolerance categories: Clear water, Silted/murky water, No preference |
| Tolerance |
pH tolerance - Acidic |
pH tolerance - Acidic |
| Tolerance |
pH tolerance - Alkaline |
pH tolerance - Alkaline |
| Tolerance |
pH tolerance - Intermediate |
pH tolerance - Intermediate |
| Waterbody Type |
Brackish waters |
taxon is found in brackish waters |
| Waterbody Type |
Cold springs |
taxon is found in cold springs |
| Waterbody Type |
Ephemeral lotic waters |
taxon is found in ephemeral lotic waters |
| Waterbody Type |
Headwater streams |
taxon is found in headwater streams |
| Waterbody Type |
Lake |
taxon is found in lakes |
| Waterbody Type |
Other |
taxon is found in an unlisted waterbody type, |
| Waterbody Type |
Pond |
taxon is found in ponds |
| Waterbody Type |
Primary waterbody |
Primary waterbody type where organism is found |
| Waterbody Type |
River |
taxon is found in rivers |
| Waterbody Type |
Second-fourth order streams |
taxon is found in second-fourth order streams |
| Waterbody Type |
Temporary lentic waters |
taxon is found in temporary lentic waters |
| Waterbody Type |
Warm springs |
taxon is found in warm springs |
| Waterbody Type |
Wetlands |
taxon is found in wetlands |
